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The beagle is a
breed A breed is a specific group of domestic animals having homogeneous appearance (phenotype), homogeneous behavior, and/or other characteristics that distinguish it from other organisms of the same species. In literature, there exist several slig ...
of small
scent hound Franz Rudolf Frisching in the uniform of an officer of the Bernese Huntsmen Corps with his Berner Laufhund, painted by Jean Preudhomme in 1785 Scent hounds (or scenthounds) are a Dog type, type of hound that primarily hunts by scent rather than ...
, similar in appearance to the much larger
foxhound A foxhound is a Dog type, type of large hunting hound bred for strong hunting instincts, a keen sense of smell, and their bark, energy, drive, and speed. In fox hunting, the foxhound's namesake, packs of foxhounds track quarry, followed—usuall ...
. The beagle was developed primarily for
hunting Hunting is the human activity, human practice of seeking, pursuing, capturing, or killing wildlife or feral animals. The most common reasons for humans to hunt are to harvest food (i.e. meat) and useful animal products (fur/hide (skin), hide, ...
hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores, and live solitarily or in pairs. They nest in slight depressions called forms, and their young are able to fend for themselves shortly after birth. The g ...
, known as beagling. Possessing a great sense of smell and superior tracking instincts, the beagle is the primary breed used as a
detection dog A detection dog or sniffer dog is a dog that is trained to use its senses to detect substances such as explosives, illegal drugs, wildlife scat, currency, blood, and contraband electronics such as illicit mobile phones. The sense most used ...
for prohibited agricultural imports and foodstuffs in
quarantine A quarantine is a restriction on the movement of people, animals and goods which is intended to prevent the spread of disease or pests. It is often used in connection to disease and illness, preventing the movement of those who may have been ...
around the world. The beagle is intelligent and is a popular
pet A pet, or companion animal, is an animal kept primarily for a person's company or entertainment rather than as a working animal, livestock, or a laboratory animal. Popular pets are often considered to have attractive appearances, intelligence ...
due to its size, good temper, and a lack of inherited health problems. The modern breed was developed in Great Britain around the 1830s from several breeds, including the Talbot Hound, the
North Country Beagle The North Country Beagle, Northern Hound or Northern Beagle was a breed of dog that existed in Britain probably until early in the 19th century. The exact date of its extinction is not known; it is likely that it was gradually interbred with othe ...
, the Southern Hound, and possibly the Harrier. Beagles have been depicted in popular culture since Elizabethan times in literature and paintings and more recently in film, television, and comic books.


History

The origin of the beagle is not known. In the 11th century,
William the Conqueror William I; ang, WillelmI (Bates ''William the Conqueror'' p. 33– 9 September 1087), usually known as William the Conqueror and sometimes William the Bastard, was the first House of Normandy, Norman List of English monarchs#House of Norman ...
brought the St. Hubert Hound and the Talbot hound to Britain. In Britain, both of these strains were then crossed with
Greyhound The English Greyhound, or simply the Greyhound, is a breed of dog, a sighthound which has been bred for coursing, greyhound racing and hunting. Since the rise in large-scale adoption of retired racing Greyhounds, the breed has seen a resurgenc ...
s to give them speed and stamina for deer hunting. Beagles are similar to the Harrier and the extinct Southern Hound, though they are smaller and slower. From medieval times, ''beagle'' was used as a generic description for the smaller hounds, though these dogs differed considerably from the modern breed. Miniature breeds of beagle-type dogs were known from the times of Edward II and Henry VII, who both had packs of Glove Beagles, so named since they were small enough to fit on a glove, and
Queen Elizabeth I Elizabeth I (7 September 153324 March 1603) was Queen of England and Ireland from 17 November 1558 until her death in 1603. Elizabeth was the last of the five House of Tudor monarchs and is sometimes referred to as the "Virgin Queen". El ...
kept a breed known as a Pocket Beagle, which stood at the shoulder. Small enough to fit in a "pocket" or saddlebag, they rode along on the hunt. The larger hounds would run the prey to ground, then the hunters would release the small dogs to continue the chase through underbrush. Elizabeth I referred to the dogs as her ''singing beagles'' and often entertained guests at her royal table by letting her Pocket Beagles cavort amid their plates and cups. 19th-century sources refer to these breeds interchangeably and it is possible that the two names refer to the same small variety. In George Jesse's ''Researches into the History of the British Dog'' from 1866, the early 17th-century poet and writer
Gervase Markham Gervase (or Jervis) Markham (ca. 1568 – 3 February 1637) was an English poet and writer. He was best known for his work '' The English Huswife, Containing the Inward and Outward Virtues Which Ought to Be in a Complete Woman'', first publishe ...
is quoted referring to the beagle as small enough to sit on a man's hand and to the: By the 18th century, two breeds had been developed for hunting hare and rabbit: the Southern Hound and the
North Country Beagle The North Country Beagle, Northern Hound or Northern Beagle was a breed of dog that existed in Britain probably until early in the 19th century. The exact date of its extinction is not known; it is likely that it was gradually interbred with othe ...
(or Northern Hound). The Southern Hound, a tall, heavy dog with a square head, and long, soft ears, was common from south of the
River Trent The Trent is the third-longest river in the United Kingdom. Its source is in Staffordshire, on the southern edge of Biddulph Moor. It flows through and drains the North Midlands. The river is known for dramatic flooding after storms and ...
and probably closely related to the Talbot Hound. Though slow, it had stamina and an excellent scenting ability. The North Country Beagle, possibly a cross between an offshoot of the Talbot stock and a Greyhound, was bred chiefly in
Yorkshire Yorkshire ( ; abbreviated Yorks), formally known as the County of York, is a historic county in northern England and by far the largest in the United Kingdom. Because of its large area in comparison with other English counties, functions have ...
and was common in the northern counties. It was smaller than the Southern Hound, less heavy-set, and with a more pointed muzzle. It was faster than its southern counterpart but its scenting abilities were less well-developed. Standards for the Pocket Beagle were drawn up as late as 1901; these genetic lines are now extinct, although modern breeders have attempted to recreate the variety.


Development of the modern breed

Reverend Phillip Honeywood established a beagle pack in
Essex Essex () is a county in the East of England. One of the home counties, it borders Suffolk and Cambridgeshire to the north, the North Sea to the east, Hertfordshire to the west, Kent across the estuary of the River Thames to the south, and Grea ...
in the 1830s and it is believed that this pack formed the basis for the modern breed. Although details of the pack's lineage are not recorded, it is thought that North Country Beagles and Southern Hounds were strongly represented;
William Youatt William Youatt (1776 – 5 February 1847) was an English veterinary surgeon and animal welfare writer. Life Youatt was the son of a non-conformist minister. He was educated for the nonconformist ministry, and undertook ministerial and scholastic ...
suspected that Harriers formed a good majority of the beagle's bloodline, but the origin of the Harrier is itself obscure. Honeywood's Beagles were small, standing at about at the shoulder, and pure white according to John Mills (writing in ''The Sportsman's Library'' in 1845). Prince Albert and Lord Winterton also had Beagle packs around this time, and royal favor no doubt led to some revival of interest in the breed, but Honeywood's pack was regarded as the finest of the three. Although credited with the development of the modern breed, Honeywood concentrated on producing dogs for hunting and it was left to Thomas Johnson to refine the breeding to produce dogs that were both attractive and capable hunters. Two strains were developed: the rough-coated and smooth-coated varieties. The rough-coated beagle survived until the beginning of the 20th century, and there were even records of one making an appearance at a dog show as late as 1969, but this variety is now extinct, having probably been absorbed into the standard beagle bloodline. In the 1840s, a standard beagle type was beginning to develop; the distinction between the North Country Beagle and Southern Hound had been lost, but there was still a large variation in size, character, and reliability among the emerging packs. In 1856, "Stonehenge" (the pseudonym of
John Henry Walsh John Henry Walsh FRCS (21 October 1810 – 12 February 1888) was an English sports writer born in Hackney, London who wrote under the pseudonym "Stonehenge." Walsh was educated in private schools and became a fellow of the Royal College of Surge ...
), writing in the ''Manual of British Rural Sports'', was still dividing beagles into four varieties: the medium beagle; the dwarf or lapdog beagle; the fox beagle (a smaller, slower version of the Foxhound); and the rough-coated or terrier beagle, which he classified as a cross between any of the other varieties and one of the Scottish
terrier Terrier (from Latin ''terra'', 'earth') is a type of dog originally bred to hunt vermin. A terrier is a dog of any one of many breeds or landraces of the terrier type, which are typically small, wiry, game, and fearless. Terrier breeds vary ...
breeds. Stonehenge also gives the start of a standard description: By 1887 the threat of extinction was on the wane: there were 18 beagle packs in England. The Beagle Club was formed in 1890 and the first standard drawn up at the same time. The following year the Association of Masters of Harriers and Beagles was formed. Both organisations aimed to further the best interests of the breed, and both were keen to produce a standard type of beagle. By 1902, the number of packs had risen to 44.


Export

Beagles were in the United States by the 1840s at the latest, but the first dogs were imported strictly for hunting and were of variable quality. Since Honeywood had only started breeding in the 1830s, it is unlikely these dogs were representative of the modern breed, and the description of them as looking like straight-legged Dachshunds with weak heads has little resemblance to the standard. Serious attempts at establishing a quality bloodline began in the early 1870s when General Richard Rowett from
Illinois Illinois ( ) is a state in the Midwestern United States. Its largest metropolitan areas include the Chicago metropolitan area, and the Metro East section, of Greater St. Louis. Other smaller metropolitan areas include, Peoria and Rockf ...
imported some dogs from England and began breeding. Rowett's Beagles are believed to have formed the models for the first American standard, drawn up by Rowett, L. H. Twadell, and Norman Ellmore in 1887. The beagle was accepted as a breed by the American Kennel Club (AKC) in 1885. In the 20th century the breed has spread worldwide.


Popularity

On its formation, the Association of Masters of Harriers and Beagles took over the running of a regular show at
Peterborough Peterborough () is a cathedral city in Cambridgeshire, east of England. It is the largest part of the City of Peterborough unitary authority district (which covers a larger area than Peterborough itself). It was part of Northamptonshire until ...
that had started in 1889, and the Beagle Club in the UK held its first show in 1896. The regular showing of the breed led to the development of a uniform type, and the beagle continued to prove a success up until the outbreak of
World War I World War I (28 July 1914 11 November 1918), often abbreviated as WWI, was one of the deadliest global conflicts in history. Belligerents included much of Europe, the Russian Empire, the United States, and the Ottoman Empire, with fightin ...
when all shows were suspended. After the war, the breed was again struggling for survival in the UK: the last of the Pocket Beagles was probably lost during this time, and registrations fell to an all-time low. A few breeders (notably Reynalton Kennels) managed to revive interest in the dog and by
World War II World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the vast majority of the world's countries—including all of the great powers—forming two opposing ...
, the breed was once again doing well. Registrations dropped again after the end of the war but almost immediately recovered. As purebred dogs, beagles have always been more popular in the United States and Canada than in their native country England. The National Beagle Club of America was formed in 1888 and by 1901 a beagle had won a Best in Show title. As in the UK, activity during World War I was minimal, but the breed showed a much stronger revival in the U.S. when hostilities ceased. In 1928 it won a number of prizes at the
Westminster Kennel Club The Westminster Kennel Club Dog Show is an all-breed conformation show, traditionally held annually at New York City's Madison Square Garden. The Westminster Kennel Club Dog Show is one of a handful of benched shows in the United States. Dogs ...
's show and by 1939 a beagle – Champion Meadowlark Draughtsman – had captured the title of top-winning American-bred dog for the year. On 12 February 2008, a beagle, K-Run's Park Me In First (Uno), won the Best In Show category at the Westminster Kennel Club show for the first time in the competition's history. In North America they have been consistently in the top-ten most-popular breeds for over 30 years. From 1953 to 1959 the beagle was ranked No. 1 on the list of the American Kennel Club's registered breeds; in 2005 and 2006 it ranked 5th out of the 155 breeds registered. In the UK they are not quite so popular, placing 28th and 30th in the rankings of registrations with the Kennel Club in 2005 and 2006 respectively. In the United States the beagle ranked 4th most popular breed in 2012 and 2013, behind the Labrador Retriever (#1),
German Shepherd The German Shepherd or Alsatian is a German breed of working dog of medium to large size. The breed was developed by Max von Stephanitz using various traditional German herding dogs from 1899. It was originally bred as a herding dog, for ...
(#2), and
Golden Retriever The Golden Retriever is a Scottish breed of retriever dog of medium size. It is characterised by a gentle and affectionate nature and a striking golden coat. It is commonly kept as a pet and is among the most frequently registered breeds ...
(#3) breeds.


Name

According to the ''
Oxford English Dictionary The ''Oxford English Dictionary'' (''OED'') is the first and foundational historical dictionary of the English language, published by Oxford University Press (OUP). It traces the historical development of the English language, providing a co ...
'', the first mention of the beagle by name in English literature dates from c. 1475 in '' The Squire of Low Degree''. The origin of the word "beagle" is uncertain, although it has been suggested that the word derives from the French ''begueule'' which means "gate throat". It is not known why the black and tan Kerry Beagle, present in Ireland since Celtic times, has the ''beagle'' description, since at it is significantly taller than the modern day beagle, and in earlier times was even larger. Some writers suggest that the beagle's scenting ability may have come from cross-breeding earlier strains with the Kerry Beagle. Originally used for hunting stags, it is today used for hare and
drag hunting Drag hunting or draghunting is a form of equestrian sport, where mounted riders hunt the trail of an artificially laid scent with hounds. Description Drag hunting is conducted in a similar manner to fox hunting, with a field of mounted riders fol ...
.


Appearance

The general appearance of the beagle resembles a miniature
Foxhound A foxhound is a Dog type, type of large hunting hound bred for strong hunting instincts, a keen sense of smell, and their bark, energy, drive, and speed. In fox hunting, the foxhound's namesake, packs of foxhounds track quarry, followed—usuall ...
, but the head is broader and the muzzle shorter, the expression completely different and the legs shorter in proportion to the body. They are generally between high at the
withers The withers is the ridge between the shoulder blades of an animal, typically a quadruped. In many species, it is the tallest point of the body. In horses and dogs, it is the standard place to measure the animal's height. In contrast, cattle a ...
and weigh between , with females being slightly smaller than males on average. They have a smooth, somewhat domed
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
with a medium-length, square-cut muzzle, and a black (or occasionally liver) gumdrop nose. The jaw is strong, and the teeth scissor together with the upper teeth fitting perfectly over the lower teeth. Both sets align square to the jaw. The eyes are large, hazel or brown, with a mild, hound-like pleading look. The large ears are long, soft, and low-set, turning towards the cheeks slightly and rounded at the tips. Beagles have a strong, medium-length neck (which is long enough for them to easily bend to the ground to pick up a scent), with little folding in the skin but some evidence of a
dewlap A dewlap is a longitudinal flap of skin or similar flesh that hangs beneath the lower jaw or neck of many vertebrates. More loosely, it can be various similar structures in the neck area, such as those caused by a double chin or the submandibul ...
; a broad chest narrowing to a tapered
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the to ...
and waist and a long, slightly curved tail (known as the "stern") tipped with white. The white tip, known as the flag, was bred for selectively, as the tail remains easily seen when the dog's head is down following a scent. The tail does not curl over the back, but is held upright when the dog is active. The beagle has a muscular body and a medium-length, smooth, hard coat. The front legs are straight and carried under the body while the rear legs are muscular and well bent at the stifles. The tricolored beaglewhite with large black areas and light brown shadingis the most common. Tricolored beagles occur in a number of shades, from the "Classic Tri" with a jet black saddle (also known as "Blackback"), to the "Dark Tri" (where faint brown markings are intermingled with more prominent black markings), to the "Faded Tri" (where faint black markings are intermingled with more prominent brown markings). Some tricolored dogs have a broken pattern, sometimes referred to as ''pied''. These dogs have mostly white coats with patches of black and brown hair. Tricolor beagles are almost always born black and white. The white areas are typically set by eight weeks, but the black areas may fade to brown as the puppy matures. (The brown may take between one and two years to fully develop.) Some beagles gradually change colour during their lives, and may lose their black markings entirely. Two-colour varieties always have a white base colour with areas of the second colour. Tan and white is the most common two-colour variety, but there is a wide range of other colours including lemon, a very light tan; red, a reddish, almost orange, brown; and liver, a darker brown, and black. Liver is not common and is not permitted in some standards; it tends to occur with yellow eyes. Ticked or mottled varieties may be either white or black with different coloured flecks (''ticking''), such as the blue-mottled or bluetick beagle, which has spots that appear to be a midnight-blue colour, similar to the colouring of the
Bluetick Coonhound The Bluetick Coonhound is a breed of coonhound originating in the United States. The Bluetick Coonhound is known for its friendly personality, cold nose, and deep bawl mouth. It is most commonly used as a raccoon hunting dog, but may also be kep ...
. Some tricolour beagles also have ticking of various colours in their white areas.


Sense of smell

Alongside the
Bloodhound The bloodhound is a large scent hound, originally bred for hunting deer, wild boar and, since the Middle Ages, for tracking people. Believed to be descended from hounds once kept at the Abbey of Saint-Hubert, Belgium, in French it is called, ...
and
Basset Hound The Basset Hound is a short-legged breed of dog in the hound family. The Basset is a scent hound that was originally bred for the purpose of hunting hare. Their sense of smell and ability to ''ground-scent'' is second only to the Bloodhound.Har ...
, the beagle has one of the best developed senses of smell of any dog. In the 1950s, John Paul Scott and John Fuller began a 13-year study of canine behavior. As part of this research, they tested the scenting abilities of various breeds by putting a mouse in a one-acre field and timing how long it took the dogs to find it. The beagles found it in less than a minute while
Fox Terrier Fox Terriers are two different breeds of the terrier dog type: the Smooth Fox Terrier and the Wire Fox Terrier. Both of these breeds originated in the 19th century from a handful of dogs who are descended from earlier varieties of British terr ...
s took 15 minutes and
Scottish Terrier The Scottish Terrier ( gd, Abhag Albannach; also known as the Aberdeen Terrier), popularly called the Scottie, is a breed of dog. Initially one of the highland breeds of terrier that were grouped under the name of ''Skye Terrier'', it is one ...
s failed to find it at all. The long ears and large lips of the beagle probably assist in trapping the scents close to the nose.


Variations


Breed varieties

The American Kennel Club recognises two separate varieties of beagle: the 13-inch for hounds less than , and the 15-inch for those between . The Canadian Kennel Club recognises a single type, with a height not exceeding . The Kennel Club (UK) and FCI affiliated clubs recognise a single type, with a height of between . English and American varieties are sometimes mentioned. However, there is no official recognition from any Kennel Club for this distinction. Beagles fitting the American Kennel Club standard – which disallows animals over – are smaller on average than those fitting the Kennel Club standard which allows heights up to . Pocket Beagles are sometimes advertised for sale but while the UK Kennel Club originally specified a standard for the Pocket Beagle in 1901, the variety is now not recognised by any Kennel Club. A strain known as Patch Hounds was developed by Willet Randall and his family from 1896 specifically for their rabbit hunting ability. They trace their bloodline back to Field Champion Patch, but do not necessarily have a patchwork marking.


Crossbreeds

In the 1850s,
John Henry Walsh John Henry Walsh FRCS (21 October 1810 – 12 February 1888) was an English sports writer born in Hackney, London who wrote under the pseudonym "Stonehenge." Walsh was educated in private schools and became a fellow of the Royal College of Surge ...
(Stonehenge) recommended a cross between a Beagle and a
Scottish Terrier The Scottish Terrier ( gd, Abhag Albannach; also known as the Aberdeen Terrier), popularly called the Scottie, is a breed of dog. Initially one of the highland breeds of terrier that were grouped under the name of ''Skye Terrier'', it is one ...
as a retriever. He found the
crossbreed A crossbreed is an organism with purebred parents of two different breeds, varieties, or populations. ''Crossbreeding'', sometimes called "designer crossbreeding", is the process of breeding such an organism, While crossbreeding is used to mai ...
to be a good worker, silent and obedient, but it had the drawback that it was small and could barely carry a hare. More recently, the trend has been for " designer dogs" and one of the most popular has been the Beagle/
Pug The Pug is a breed of dog originally from China, with physically distinctive features of a wrinkly, short-muzzled face and curled tail. The breed has a fine, glossy coat that comes in a variety of colors, most often light brown (Fawn (colour) ...
cross known as a Puggle. Some puppies of this cross are less excitable than a Beagle and with a lower exercise requirement, similar to the Pug parent; but many are highly excitable and require vigorous exercise.


Temperament

The beagle has an even temper and gentle disposition. Described in several breed standards as "merry", they are amiable and typically neither aggressive nor timid, although this depends on the individual. They enjoy company, and although they may initially be standoffish with strangers, they are easily won over. They make poor guard dogs for this reason, although their tendency to bark or
howl Howl most often refers to: *Howling, an animal vocalization in many canine species *Howl (poem), a 1956 poem by Allen Ginsberg Howl may also refer to: Film * ''The Howl'', a 1970 Italian film * ''Howl'' (2010 film), a 2010 American arthouse b ...
when confronted with the unfamiliar makes them good watch dogs. In a 1985 study conducted by Ben and Lynette Hart, the beagle was given the highest excitability rating, along with the
Yorkshire Terrier The Yorkshire Terrier (often shortened as Yorkie) is one of the smallest dog breeds of the terrier type and indeed of any dog breed. The breed developed during the 19th century in Yorkshire, England.Cairn Terrier The Cairn Terrier is a terrier breed originating in the Scottish Highlands and recognized as one of Scotland's earliest working dogs. The breed was given the name Cairn because the breed's function was to hunt and chase quarry between the ca ...
,
Miniature Schnauzer The Miniature Schnauzer is a breed of small dog of the Schnauzer type that originated in Germany in the mid-to-late 19th century. Miniature Schnauzers may have been developed from the smallest specimens of the Standard Schnauzer, or crosses be ...
,
West Highland White Terrier The West Highland White Terrier, commonly known as the Westie, is a breed of dog from Scotland with a distinctive white harsh coat with a somewhat soft white undercoat. It is a medium-sized terrier, although with longer legs than other Scot ...
, and
Fox Terrier Fox Terriers are two different breeds of the terrier dog type: the Smooth Fox Terrier and the Wire Fox Terrier. Both of these breeds originated in the 19th century from a handful of dogs who are descended from earlier varieties of British terr ...
. They are ranked 72nd in
Stanley Coren Stanley Coren (born 1942) is a psychology professor, neuropsychological researcher and writer on the intelligence, mental abilities and history of dogs. He works in research and instructs in psychology at the University of British Columbia in V ...
's ''
The Intelligence of Dogs ''The Intelligence of Dogs'' is a 1994 book on dog intelligence by Stanley Coren, a professor of canine psychology at the University of British Columbia. The book explains Coren's theories about the differences in intelligence between various ...
'', as Coren places them among the group with the lowest degree of working/obedience intelligence. Coren's scale, however, does not assess understanding, independence, or creativity. Beagles are excellent with children and this is one of the reasons they have become popular family pets. Beagles are pack animals; they are prone to
separation anxiety Separation anxiety disorder (SAD) is an anxiety disorder in which an individual experiences excessive anxiety regarding separation from home and/or from people to whom the individual has a strong emotional attachment (e.g., a parent, caregiver, ...
, a condition which causes them to destroy things when left unattended. Not all beagles will howl, but most will bark when confronted with strange situations, and some will bay (also referred to as "speaking", "giving tongue", or "opening") when they catch the scent of potential quarry. They also generally get along well with cats and other dogs. They are not too demanding with regard to exercise; their inbred stamina means they do not easily tire when exercised, but they also do not need to be worked to exhaustion before they will rest. Regular exercise helps ward off the weight gain to which the breed is prone.


Health

The typical longevity of beagles is 12–15 years, which is a common lifespan for dogs of their size. Beagles may be prone to
epilepsy Epilepsy is a group of non-communicable neurological disorders characterized by recurrent epileptic seizures. Epileptic seizures can vary from brief and nearly undetectable periods to long periods of vigorous shaking due to abnormal electrica ...
, but this can often be controlled with medication. Hypothyroidism and a number of types of dwarfism occur in beagles. Two conditions in particular are unique to the breed: "Funny Puppy", in which the puppy is slow to develop and eventually develops weak legs, a crooked back and although normally healthy, is prone to a range of illnesses; and Musladin-Lueke syndrome (MLS) in which the eyes are slanted and the outer toes are underdeveloped but otherwise development is as normal.
Hip dysplasia Hip dysplasia is an abnormality of the hip joint where the socket portion does not fully cover the ball portion, resulting in an increased risk for joint dislocation. Hip dysplasia may occur at birth or develop in early life. Regardless, it doe ...
, common in Harriers and in some larger breeds, is rarely considered a problem in beagles. Beagles are considered a chondrodystrophic breed, meaning that they are prone to types of disk diseases. In rare cases, beagles may develop immune mediated polygenic arthritis (where the immune system attacks the joints) even at a young age. The symptoms can sometimes be relieved by steroid treatments. Another rare disease in the breed is neonatal cerebellar cortical degeneration. Affected puppies are slow, have lower co-ordination, fall more often, and do not have a normal gait. It has an estimated carrier rate of 5% and affected rate of 0.1%. A genetic test is available. Their long floppy ears can mean that the inner ear does not receive a substantial air flow or that moist air becomes trapped, and this can lead to ear infections. Beagles may also be affected by a range of eye problems; two common ophthalmic conditions in beagles are
glaucoma Glaucoma is a group of eye diseases that result in damage to the optic nerve (or retina) and cause vision loss. The most common type is open-angle (wide angle, chronic simple) glaucoma, in which the drainage angle for fluid within the eye rem ...
and
corneal dystrophy Corneal dystrophy is a group of rare hereditary disorders characterised by bilateral abnormal deposition of substances in the transparent front part of the eye called the cornea. Signs and symptoms Corneal dystrophy may not significantly affect v ...
. "
Cherry eye Cherry eye is a disorder of the nictitating membrane (NM), also called the third eyelid, present in the eyes of dogs and cats.Gelatt, K. N. (2000). ''Essentials of Veterinary Ophthalmology''. Baltimore: Lippincott Williams & Wilkins. Cherry eye ...
", a prolapse of the gland of the
third eyelid The nictitating membrane (from Latin '' nictare'', to blink) is a transparent or translucent third eyelid present in some animals that can be drawn across the eye from the medial canthus to protect and moisten it while maintaining vision. All ...
, and
distichiasis A distichia is an eyelash that arises from an abnormal part of the eyelid. This abnormality, attributed to a genetic mutation, is known to affect dogs and humans. Distichiae usually exit from the duct of the meibomian gland at the eyelid margin. T ...
, a condition in which eyelashes grow into the eye causing irritation, sometimes exist; both these conditions can be corrected with surgery. They can suffer from several types of retinal atrophy. Failure of the nasolacrimal drainage system can cause dry eye or leakage of tears onto the face. As field dogs they are prone to minor injuries such as cuts and sprains, and, if inactive, obesity is a common problem as they will eat whenever food is available and rely on their owners to regulate their weight. When working or running free they are also likely to pick up parasites such as
flea Flea, the common name for the order Siphonaptera, includes 2,500 species of small flightless insects that live as external parasites of mammals and birds. Fleas live by ingesting the blood of their hosts. Adult fleas grow to about long, a ...
s,
tick Ticks (order Ixodida) are parasitic arachnids that are part of the mite superorder Parasitiformes. Adult ticks are approximately 3 to 5 mm in length depending on age, sex, species, and "fullness". Ticks are external parasites, living by ...
s, harvest mites, and
tapeworm Eucestoda, commonly referred to as tapeworms, is the larger of the two subclasses of flatworms in the class Cestoda (the other subclass is Cestodaria). Larvae have six posterior hooks on the scolex (head), in contrast to the ten-hooked Cesto ...
s, and irritants such as grass seeds can become trapped in their eyes, soft ears, or paws. Beagles may exhibit a behavior known as reverse sneezing, in which they sound as if they are choking or gasping for breath, but are actually drawing air in through the mouth and nose. The exact cause of this behavior is not known, but it can be a common occurrence and is not harmful to the dog.


Birth and Reproduction

The average size of a beagle litter is six puppies. When mother beagles give birth to litters of puppies, the little pups weigh just a few ounces each.


Hunting

Beagles were developed primarily for hunting hare, an activity known as beagling. They were seen as ideal hunting companions for the elderly who could follow on horseback without exerting themselves, for young hunters who could keep up with them on ponies, and for the poorer hunters who could not afford to maintain a stable of good hunting horses. Before the advent of the fashion for foxhunting in the 19th century, hunting was an all day event where the enjoyment was derived from the chase rather than the kill. In this setting the tiny beagle was well matched to the hare, as unlike Harriers they would not quickly finish the hunt, but because of their excellent scent-tracking skills and stamina they were almost guaranteed to eventually catch the hare. The beagle packs would run closely together ("so close that they might be covered with a sheet") which was useful in a long hunt, as it prevented stray dogs from obscuring the trail. In thick undergrowth they were also preferred to
spaniel A spaniel is a type of gun dog. Spaniels were especially bred to flush game out of denser brush. By the late 17th century, spaniels had been specialized into water and land breeds. The extinct English Water Spaniel was used to retrieve water ...
s when hunting
pheasant Pheasants ( ) are birds of several genera within the family Phasianidae in the order Galliformes. Although they can be found all over the world in introduced (and captive) populations, the pheasant genera native range is restricted to Eurasia ...
. With the fashion for faster hunts, the beagle fell out of favor for chasing hare, but was still employed for rabbit hunting. In ''Anecdotes of Dogs'' (1846), Edward Jesse says: In the United States they appear to have been employed chiefly for hunting rabbits from the earliest imports. Hunting hare with beagles became popular again in Britain in the mid-19th century and continued until it was made illegal in Scotland by the Protection of Wild Mammals (Scotland) Act 2002 and in England and Wales by the
Hunting Act 2004 The Hunting Act 2004 (c 37) is an Act of Parliament, Act of the Parliament of the United Kingdom which bans the hunting of most wild mammals (notably Red fox, foxes, deer, European hare, hares and American mink, mink) with dogs in England and W ...
. Under this legislation beagles may still pursue rabbits with the landowner's permission.
Drag hunting Drag hunting or draghunting is a form of equestrian sport, where mounted riders hunt the trail of an artificially laid scent with hounds. Description Drag hunting is conducted in a similar manner to fox hunting, with a field of mounted riders fol ...
is popular where hunting is no longer permitted or for those owners who do not wish to participate in hunting a live animal, but still wish to exercise their dog's innate skills. The traditional foot pack consists of up to 40 beagles, marshaled by a Huntsman who directs the pack and who is assisted by a variable number of whippers-in whose job is to return straying hounds to the pack. The Master of the Hunt is in overall day-to-day charge of the pack, and may or may not take on the role of Huntsman on the day of the hunt. As hunting with beagles was seen as ideal for young people, many of the British public schools traditionally maintained beagle packs. Protests were lodged against Eton's use of beagles for hunting as early as 1902 but the pack is still in existence today. In 2001, the Wye College beagle pack was taken by the
Animal Liberation Front The Animal Liberation Front (ALF) is an international, leaderless, decentralized political and social resistance movement that engages in and promotes non-violent direct action in protest against incidents of animal cruelty. It originated in th ...
. School and university packs are still maintained by Eton, Marlborough,
Radley Radley is a village and civil parish about northeast of the centre of Abingdon, Oxfordshire. The parish includes the hamlet of Lower Radley on the River Thames. It was part of Berkshire until the 1974 boundary changes transferred it to Oxfor ...
, the
Royal Agricultural University ;(from Virgil's Georgics)"Caring for the Fieldsand the Beasts" , established = 2013 - University status – College , type = Public , president = King Charles , vice_chancellor = Peter McCaffery , students ...
, and Christ Church, Oxford. In addition to organised beagling, beagles have been used for hunting or flushing to guns (often in pairs) a wide range of game including
snowshoe hare The snowshoe hare (''Lepus americanus''), also called the varying hare or snowshoe rabbit, is a species of hare found in North America. It has the name "snowshoe" because of the large size of its hind feet. The animal's feet prevent it from sin ...
,
cottontail rabbit Cottontail rabbits are the leporid species in the genus ''Sylvilagus'', found in the Americas. Most ''Sylvilagus'' species have stub tails with white undersides that show when they retreat, giving them their characteristic name. However, this ...
s,
game bird Galliformes is an order of heavy-bodied ground-feeding birds that includes turkeys, chickens, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed dispersers and predators, and are ofte ...
s,
roe deer The roe deer (''Capreolus capreolus''), also known as the roe, western roe deer, or European roe, is a species of deer. The male of the species is sometimes referred to as a roebuck. The roe is a small deer, reddish and grey-brown, and well-adapt ...
,
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of we ...
, bobcat, coyote,
wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species is ...
, and foxes, and have even been recorded as being used to hunt
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
. In most of these cases, the beagle is employed as a gun dog, flushing game for hunter's guns.


Detection

Beagles are used as detection dogs in the Beagle Brigade of the
United States Department of Agriculture The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of com ...
. These dogs are used to detect food items in luggage being taken into the United States. After trialling several breeds, beagles were chosen because they are relatively small and unintimidating for people who are uncomfortable around dogs, easy to care for, intelligent and work well for rewards. They are also used for this purpose in a number of other countries including by the Ministry of Agriculture and Forestry in New Zealand, the
Australian Quarantine and Inspection Service The Australian Quarantine and Inspection Service was the Australian government agency responsible for enforcing Australian quarantine laws, as part of the Department of Agriculture. Following a period operating under the name DAFF Biosecurit ...
, and in Canada, Japan, and the People's Republic of China. Larger breeds are generally used for detection of explosives as this often involves climbing over luggage and on large conveyor belts, work for which the smaller Beagle is not suited.


Testing

Beagles are the dog breed most often used in
animal testing Animal testing, also known as animal experimentation, animal research, and ''in vivo'' testing, is the use of non-human animals in experiments that seek to control the variables that affect the behavior or biological system under study. This ...
, due to their size and passive nature. In the United States, as many as 65,000 beagles are used every year for medical, cosmetic, beauty, and other chemical tests. They are purpose bred and live their lives in cages undergoing experiments. The Rescue + Freedom Project (formerly Beagle Freedom Project) has successfully advocated for beagles to be released from labs. This organisation has freed hundreds of animals. Beagles are used in a range of research procedures: fundamental biological research, applied human medicine, applied veterinary medicine, and protection of man, animals, or the environment. Of the 8,018 dogs used in testing in the UK in 2004, 7,799 were beagles (97.3%). In the UK, the
Animals (Scientific Procedures) Act 1986 The Animals (Scientific Procedures) Act 1986, sometimes referred to as ASPA, is an Act of Parliament, Act of the Parliament of the United Kingdom (1986 c. 14) passed in 1986, which regulates the use of animals used for research in the UK. The Act ...
gave special status to primates, equids, cats and dogs and in 2005 the Animal Procedures Committee (set up by the act) ruled that testing on mice was preferable, even though a greater number of individual animals were involved. In 2005 beagles were involved in less than 0.3% of the total experiments on animals in the UK, but of the 7670 experiments performed on dogs 7406 involved beagles (96.6%). Most dogs are bred specifically for this purpose, by companies such as Harlan. In the UK companies breeding animals for research must be licensed under the Animals (Scientific Procedures) Act.


Bans and activism against beagle testing

Testing of cosmetic products on animals is banned in the member states of the European Community, although France protested the ban and has made efforts to have it lifted. It is permitted in the United States but is not mandatory if safety can be ascertained by other methods, and the test species is not specified by the
Food and Drug Administration The United States Food and Drug Administration (FDA or US FDA) is a List of United States federal agencies, federal agency of the United States Department of Health and Human Services, Department of Health and Human Services. The FDA is respon ...
(FDA). When testing toxicity of food additives, food contaminants, and some drugs and chemicals the FDA uses beagles and
miniature pig Miniature Pigs, also called mini pig, or Pygmy pig, are small breeds of domestic pig, such as the Vietnamese Pot-Bellied pig, Göttingen minipig, Juliana pig, Choctaw hog, or Kunekune (and specimens derived by crossbreeding these breeds). Mi ...
s as surrogates for direct human testing. Minnesota was the first state to enact a Beagle Freedom adoption law in 2014, mandating that dogs and cats are allowed to be adopted once they have completed research testing. Anti-
vivisection Vivisection () is surgery conducted for experimental purposes on a living organism, typically animals with a central nervous system, to view living internal structure. The word is, more broadly, used as a pejorative catch-all term for Animal testi ...
groups have reported on abuse of animals inside testing facilities. In 1997 footage secretly filmed by a freelance journalist inside
Huntingdon Life Sciences Huntingdon Life Sciences (HLS) was a contract research organisation (CRO) founded in 1951 in Cambridgeshire, England. It had two laboratories in the United Kingdom and one in the United States. With over 1,600 staff, it was until 2015 the largest ...
in the UK showed staff punching and screaming at beagles. Consort Kennels, a UK-based breeder of beagles for testing, closed down in 1997 after pressure from animal rights groups. There are various examples of activists utilizing Freedom of Information Act (FOIA) requests to obtain information about taxpayer funding of animal testing. For example, the White Coat Waste Project, a group of activists that hold that taxpayers should not have to pay $20 billion every year for experiments on animals, highlighted that the
National Institute of Allergy and Infectious Diseases The National Institute of Allergy and Infectious Diseases (NIAID, ) is one of the 27 institutes and centers that make up the National Institutes of Health (NIH), an agency of the United States Department of Health and Human Services (HHS). NIAID's ...
provided $400,000 in taxpayer money to fund experiments in which 28 beagles were infected by disease-causing parasites. The White Coat Project found reports that said dogs taking part in the experiments were “vocalizing in pain” after being injected with foreign substances. Following public outcry, People for the Ethical Treatment of Animals (PETA) made a call to action that all members of the
National Institute of Health The National Institutes of Health, commonly referred to as NIH (with each letter pronounced individually), is the primary agency of the United States government responsible for biomedical and public health research. It was founded in the late ...
resign effective immediately and that there is a "need to find a new NIH director to replace the outgoing
Francis Collins Francis Sellers Collins (born April 14, 1950) is an American physician-geneticist who discovered the genes associated with a number of diseases and led the Human Genome Project. He is the former director of the National Institutes of Health (N ...
who will shut down research that violates the dignity of nonhuman animals."


Other roles

Although bred for hunting, Beagles are versatile and are nowadays employed for various other roles in detection, therapy, and as family pets. Beagles are used as sniffer dogs for
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blatto ...
detection in Australia, and have been mentioned as possible candidates for drug and explosive detection. Because of their gentle nature and unimposing build, they are also frequently used in pet therapy, visiting the sick and elderly in hospital. In June 2006, a trained Beagle
assistance dog In general, an assistance dog, known as a service dog in the United States, is a dog trained to aid or assist an individual with a disability. Many are trained by an assistance dog organization, or by their handler, often with the help of a prof ...
was credited with saving the life of its owner after using her owner's mobile phone to dial an emergency number. In the aftermath of the
2010 Haiti earthquake A catastrophic magnitude 7.0 Mw earthquake struck Haiti at 16:53 local time (21:53 UTC) on Tuesday, 12 January 2010. The epicenter was near the town of Léogâne, Ouest department, approximately west of Port-au-Prince, Haiti's c ...
, a Beagle
search and rescue dog A search-and-rescue dog is one trained to find missing people after a natural or man-made disaster. The dogs detect human scent and have been known to find people under water, under snow, and under collapsed buildings. Applications A dog ...
with a Colombian rescue squad was credited with locating the owner of the
Hôtel Montana The Hôtel Montana is a four star luxury hotel in Haiti. It is located on Rue Frank Cardozo, Bourdon on one of the only three main roads connecting the capital to its southeastern outskirt city Pétion-Ville. It was one of the most luxurious hot ...
, who was subsequently rescued after spending 100 hours buried in the rubble. Beagles were hired by New York City to help with bedbug detection, while the role of such dogs in this type of detection may have doubts.


In popular culture

* Anthropomorphic Beagles appeared in comic strips and animated cartoons since the 1950s with
Courage the Cowardly Dog ''Courage the Cowardly Dog'' is an American animated comedy horror television series created by John R. Dilworth for Cartoon Network and distributed by Warner Bros. Domestic Television. It was produced by Dilworth's animation studio, Stretch Fil ...
and the ''
Peanuts ''Peanuts'' is a syndicated daily and Sunday American comic strip written and illustrated by Charles M. Schulz. The strip's original run extended from 1950 to 2000, continuing in reruns afterward. ''Peanuts'' is among the most popular and infl ...
'' character
Snoopy Snoopy is an anthropomorphic beagle in the comic strip ''Peanuts'' by Charles M. Schulz. He can also be found in all of the ''Peanuts'' films and television specials. Since his debut on October 4, 1950, Snoopy has become one of the most recog ...
, who was billed as "the world's most famous Beagle". *
Porthos Porthos, Baron du Vallon de Bracieux de Pierrefonds is a fictional character in the novels ''The Three Musketeers'' (1844), '' Twenty Years After'' (1845), and '' The Vicomte de Bragelonne'' (1847-1850) by Alexandre Dumas, père. He and the othe ...
is a beagle that belongs to
Jonathan Archer Jonathan Archer is a fictional character in the ''Star Trek'' franchise. He is the protagonist of the television series ''Star Trek: Enterprise'', where he is portrayed by Scott Bakula. Archer is the commanding officer of the first starship ' ...
, captain of the
Star Trek: Enterprise ''Star Trek: Enterprise'', titled simply ''Enterprise'' for its first two seasons, is an American science fiction television series created by Rick Berman and Brannon Braga. It originally aired from September 26, 2001, to May 13, 2005 on Uni ...
; *Former US President
Lyndon Baines Johnson Lyndon Baines Johnson (; August 27, 1908January 22, 1973), often referred to by his initials LBJ, was an American politician who served as the 36th president of the United States from 1963 to 1969. He had previously served as the 37th vice ...
had several beagles, and caused an outcry when he picked up one of them by its ears during an official greeting on the White House lawn. *The ship on which
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended ...
made the voyage which provided much of the inspiration for ''
On the Origin of Species ''On the Origin of Species'' (or, more completely, ''On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life''),The book's full original title was ''On the Origin of Species by Me ...
'' was named HMS ''Beagle'' after the breed, and, in turn, lent its name to the ill-fated British
Martian Mars, the fourth planet from the Sun, has appeared as a setting in works of fiction since at least the mid-1600s. It became the most popular celestial object in fiction in the late 1800s as the Moon was evidently lifeless. At the time, the pr ...
lander ''
Beagle 2 The ''Beagle 2'' is an inoperative British Mars lander that was transported by the European Space Agency's 2003 ''Mars Express'' mission. It was intended to conduct an astrobiology mission that would have looked for evidence of past life on Mar ...
''. *An American bred 15 inch male Beagle with the registered name of Ch K-Run's Park Me In First and the pet name of "Uno" won the 2008
Westminster Kennel Club Dog Show The Westminster Kennel Club Dog Show is an all-breed conformation show, traditionally held annually at New York City's Madison Square Garden. The Westminster Kennel Club Dog Show is one of a handful of benched shows in the United States. Dogs ...
. *A Canadian bred 15 inch female Beagle with the registered name of Gr Ch Tashtins Lookin For Trouble and the pet name of "Miss P" won the 2015
Westminster Kennel Club Dog Show The Westminster Kennel Club Dog Show is an all-breed conformation show, traditionally held annually at New York City's Madison Square Garden. The Westminster Kennel Club Dog Show is one of a handful of benched shows in the United States. Dogs ...
.


Notable Beagles

* Frodo, awarded the PDSA Gold Medal for animal bravery * Uno, who in 2008 became the first Beagle to win the Westminster Kennel Club Dog Show * Miss P, winner of the 2015 Westminster Kennel Club Dog Show


References

Informational notes a. In this article "Beagle" (with a capital B) is used to distinguish the modern breed from other beagle-type dogs. c. The Harts posed the following question to a panel of 96 experts, half of which were veterinary surgeons and the other half dog obedience trial judges:
d. The specific references in each of the author's works are as follows:
Shakespeare: "''Sir Toby Belch'': She's a beagle, true-bred, and one that adores me: what o' that?" ''
Twelfth Night ''Twelfth Night'', or ''What You Will'' is a romantic comedy by William Shakespeare, believed to have been written around 1601–1602 as a Twelfth Night's entertainment for the close of the Christmas season. The play centres on the twins Vi ...
'' (c. 1600) Act II Scene III
Webster: "''Mistress Tenterhook: You are a sweet beagle" '' Westward Ho'' (1607) Act III Scene IV:2
Dryden: "The rest in shape a beagle's whelp throughout, With broader forehead and a sharper snout" ''The Cock and the Fox'', and again: "About her feet were little beagles seen" in ''Palamon and Arcite'' both from '' Fables, Ancient and Modern'' (1700)
Tickell: "Here let me trace beneath the purpled morn, The deep-mouth'd beagle, and the sprightly horn" ''To a Lady before Marriage'' (published posthumously in 1749)
Fielding: "'What the devil would you have me do?' cries the Squire, turning to Blifil, 'I can no more turn her, than a beagle can turn an old hare.'" ''
The History of Tom Jones, a Foundling ''The History of Tom Jones, a Foundling'', often known simply as ''Tom Jones'', is a comic novel by English playwright and novelist Henry Fielding. It is a ''Bildungsroman'' and a picaresque novel. It was first published on 28 February 1749 in L ...
'' (1749) Chapter 7.
Cowper: "For persevering chase and headlong leaps, True beagle as the staunchest hound he keeps" ''The Progress of Error'' (1782)
Pope: "Thus on a roe the well-breath'd beagle flies, And rends his hide fresh-bleeding with the dart" ''The Iliad of Homer'' (1715–20) Book XV:697–8 Citations Bibliography * * * * * * * * * * * * * * * *


External links

* {{Authority control FCI breeds Companion dogs Scent hounds Dog breeds originating in England Hunting with hounds